tree Viburnum = (Viburnum_clemensiae,(((((((Viburnum_integrifolium,(Viburnum_japonicum,(Viburnum_lancifolium,(Viburnum_setigerum,Viburnum_phlebotrichum),((Viburnum_formosanum,Viburnum_fordiae),(Viburnum_luzonicum,Viburnum_tashiroi)),((Viburnum_erosum,Viburnum_brachyandrum),(Viburnum_anamensis,Viburnum_sempervirens)),(Viburnum_wrightii,((Viburnum_adenophorum,Viburnum_ichangense),(Viburnum_parvifolium,(Viburnum_dilatatum,(Viburnum_brevipes,Viburnum_corylifolium)))))))),(Viburnum_foetidum,(Viburnum_mullaha,((Viburnum_betulifolium,Viburnum_hupehense),(Viburnum_melanocarpum,(Viburnum_flavescens,Viburnum_lobophyllum)))))),(Viburnum_kansuense,(Viburnum_orientale,Viburnum_acerifolium))),(((Viburnum_ternatum,(Viburnum_leiocarpum,(Viburnum_inopinatum,Viburnum_sambucinum))),(Viburnum_beccarii,(Viburnum_glaberrimum,(Viburnum_hispidulum,Viburnum_vernicosum)))),(Viburnum_cylindricum,(Viburnum_coriaceum,Viburnum_hebanthum)))),((Viburnum_edule,Viburnum_koreanum),(Viburnum_trilobum,(Viburnum_opulus,Viburnum_sargentii)))),((((Viburnum_loeseneri,(Viburnum_stenocalyx,(Viburnum_ciliatum,(Viburnum_microcarpum,Viburnum_caudatum)))),(((Viburnum_jucundum,Viburnum_lautum),(Viburnum_obtusatum,(Viburnum_hartwegii,(Viburnum_costaricanum,(((Viburnum_glabratum,((Viburnum_subsessile,Viburnum_anabaptista),(Viburnum_tinoides,Viburnum_undulatum)),(Viburnum_lasiophyllum,Viburnum_jamesonii)),(Viburnum_pichinchense,Viburnum_hallii)),((Viburnum_seemenii,Viburnum_toronis,Viburnum_ayavacense),(Viburnum_triphyllum,(Viburnum_obtectum,(Viburnum_divaricatum,Viburnum_stipitatum)))))))),(Viburnum_villosum,(Viburnum_stellato_tomentosum,Viburnum_disjunctum))),(Viburnum_blandum,Viburnum_discolor,Viburnum_microphyllum,(Viburnum_venustum,(Viburnum_sulcatum,Viburnum_acutifolium))))),(Viburnum_scabrellum,(Viburnum_dentatum,Viburnum_recognitum))),((Viburnum_ellipticum,Viburnum_australe),(Viburnum_bracteatum,(Viburnum_molle,Viburnum_rafinesquianum))))),((Viburnum_davidii,(Viburnum_propinquum,(Viburnum_cinnamomifolium,(Viburnum_calvum,Viburnum_atrocyaneum)))),(Viburnum_treleasei,(Viburnum_tinus,Viburnum_rigidum)))),((((Viburnum_cotinifolium,((((Viburnum_glomeratum,Viburnum_veitchii),(Viburnum_rhytidophyllum,(Viburnum_chinshanense,Viburnum_buddleifolium))),(Viburnum_mongolicum,(Viburnum_burejaeticum,(Viburnum_lantana,Viburnum_maculatum)))),(Viburnum_macrocephalum,((Viburnum_utile,Viburnum_schensianum),(Viburnum_congestum,(Viburnum_carlesii,Viburnum_bitchiuense)))))),(Viburnum_punctatum,((Viburnum_nudum,Viburnum_cassinoides),(Viburnum_lentago,((Viburnum_elatum,Viburnum_obovatum),(Viburnum_prunifolium,Viburnum_rufidulum)))))),(Viburnum_amplificatum,(((Viburnum_subalpinum,(Viburnum_corymbiflorum,((Viburnum_suspensum,Viburnum_taitoense),(Viburnum_farreri,(Viburnum_grandiflorum,Viburnum_foetens))),((Viburnum_brevitubum,Viburnum_erubescens,Viburnum_oliganthum),(Viburnum_chingii,Viburnum_yunnanense),(Viburnum_brachybotryum,Viburnum_henryi)))),(Viburnum_sieboldii,(Viburnum_odoratissimum,Viburnum_awabuki))),((Viburnum_amplifolium,(Viburnum_pyramidatum,Viburnum_lutescens)),(Viburnum_colebrookeanum,(Viburnum_hanceanum,Viburnum_plicatum)))))),((Viburnum_urceolatum,Viburnum_taiwanianum),(Viburnum_nervosum,(Viburnum_lantanoides,(Viburnum_furcatum,Viburnum_sympodiale))))));

mrca Lentago = Viburnum_nudum, Viburnum_lentago;
mrca Tinus = Viburnum_tinus, Viburnum_davidii;
mrca Opulus = Viburnum_edule, Viburnum_opulus;
mrca Porphyrotinus = Viburnum_ellipticum, Viburnum_subsessile;
mrca Urceolata = Viburnum_urceolatum, Viburnum_taiwanianum;
mrca Pseudotinus = Viburnum_nervosum, Viburnum_furcatum;
mrca Euviburnum = Viburnum_cotinifolium, Viburnum_rhytidophyllum;
mrca Solenotinus = Viburnum_sieboldii, Viburnum_erubescens;
mrca Lutescentia = Viburnum_plicatum, Viburnum_lutescens;
mrca Mollotinus = Viburnum_ellipticum, Viburnum_molle;
mrca Oreinotinus = Viburnum_stenocalyx, Viburnum_subsessile;
mrca Succotinus = Viburnum_foetidum, Viburnum_dilatatum;
mrca Sambucina = Viburnum_beccarii, Viburnum_sambucinum;


reference = 'Donoghue (pers. comm.)';